Both engines feature a cranking system that keeps the starter engaged until complete combustion is achieved, freeing the driver from holding the key until the engine turns over. This is a nice feature, and one usually associated with expensive luxury sedans.
Both engines come with a sophisticated five-speed automatic transmission. More gears means better response for any given situation along with better efficiency and this five-speed is more flexible than a four-speed and better able to keep the engine running in its optimum rpm range, whether you're after power or fuel economy at any particular moment. The transmission is equipped with Artificial Intelligence Shift control, which changes gear-shifting patterns according to driving conditions and driver intent. It works well and seems to understand when you want to cruise and when you want to get with the program, and it shifts smoothly around town.
The 4Runner handles very well for a truck with a live rear axle. We drove V6-powered models over twisting back roads along the Oregon coast and found them easy to drive at a quick clip. We've also spent a lot of time in V8 versions around Los Angeles.
The suspension damping is excellent. When the road got bumpy, we could tell our truck had a solid rear axle rather than an independent rear suspension, but the 4Runner still handles more confidently than other live-axle SUVs. Rack-and-pinion steering gives the 4Runner quick response and good steering feel.
Still, the 4Runner is a truck, not a car. Rather than using unit-body construction like the Toyota Highlander and RAV4, the 4Runner is built on a separate ladder frame that features full-length box-section frame rails. Toyota also steered away from using an independent rear suspension like the one on the ladder-frame Ford Explorer. An independent rear suspension would have offered a cushier ride around town and allowed for a roomier interior, but off-road capability was a high priority for the 4Runner, and its live rear axle provides more suspension travel. In other words, if your driving consists almost entirely of commuting to work, hauling kids around and running errands, you might be more comfortable in a Toyota Highlander.
The 4Runner starts making a lot of sense when pull off the pavement. The ride quality on unpaved roads is smooth and well-controlled, important on long gravel treks over washboard surfaces on the way to a remote fishing spot. Well-tuned damping and progressive-rate spring bumpers are to thank here.
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